What is the meaning of Exannulate?

(a.) Having the sporangium destitute of a ring; -- said of certain genera of ferns.
Webster's Revised Unabridged Dictionary
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APAWebster's. (2011). Exannulate. Retrieved September 30, 2026, from http://smartdefine.org/exannulate/definitions/2005517
ChicagoWebster's. 2011. "Exannulate" http://smartdefine.org/exannulate/definitions/2005517 (accessed September 30, 2026).
HarvardWebster's 2011, Exannulate, Smart Define, viewed 30 September, 2026, <http://smartdefine.org/exannulate/definitions/2005517>.
MLAWebster's. "Exannulate" 27 May 2011. Web. 30 September 2026. <http://smartdefine.org/exannulate/definitions/2005517>