What is the meaning of trodden?

verb

• Put down or press the foot, place the foot.
  • usage: "For fools rush in where angels fear to tread"; "step on the brake"
  • synonyms: step, tread
• Tread or stomp heavily or roughly.
  • usage: "The soldiers trampled across the fields"
  • synonyms: tread, trample
• Crush as if by treading on.
  • usage: "tread grapes to make wine"
  • synonyms: tread
• Brace (an archer's bow) by pressing the foot against the center.
  • synonyms: tread
• Apply (the tread) to a tire.
  • synonyms: tread
• Mate with.
  • usage: "male birds tread the females"
  • synonyms: tread