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- Plant Embryology by Great Soviet Encyclopedia... layers (dermatogen, periblem, plerome), from which the permanent tissue of the adult organism develops.The Russian botanist S. M. Rozanov (1865, 1866) laid the foundation for the comparative embryology of Hydrophyllaceae, Boraginaceae, and other families. A. S. Famintsyn made a detailed study of embryogeny in both dicotyledonous and monocotyledonous plants. S. G...